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In seated pulley rows with a V-grip, I've always used the grip with the V pointing at the floor (ie. with a "underhand" grip). However, this weekend I saw a picture of Arnold demonstrating seated pulley rows with the V pointing at the roof (ie. overhand grip, if you can call it that...)
Does it make a difference?
Both work the back, underhand can use more biceps though. Same as reverse grip barbell row. overhand is the same as barbell row
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